UPDATE! - reports Reese's rep denies the split. We've put in calls to her people and we're waiting to hear back.

Reese Witherspoon, 33, and Jake Gyllenhaal, 29, have ended their relationship after two and a half years, a source tells

The couple started dating in April 2007, after meeting on the set of the film Rendition -- a CIA thriller in which they starred opposite each other.

The pair have been the target of marriage speculation almost since they went public with their relationship. Reese told Parade magazine last year: "Family is all we have in life, but I don’t know how I feel about marriage. Obviously, I'm not far enough out of being married to think about doing it again."

"I don't think about it much ... it's important to stay open to whatever your relationship evolves into. I have a certain idea of what I thought marriage is and the whole life and [the way] things are, and sometimes it doesn’t work out that way," she told the mag.

Reese split from her husband of seven years, Ryan Phillippe, in 2006. They have two children together - Ava, 10, and Deacon, 6.